%PDF-1.4
%
2 0 obj
<>
endobj
66 0 obj
<>stream
2006-06-09T09:12:33Z
page2pdf
2013-04-24T14:22:04-07:00
2013-04-24T14:22:04-07:00
PDFlib 3.02 (SunOS 5.6)
uuid:43fa354e-1dd2-11b2-0a00-4508672c7500
uuid:43fa3552-1dd2-11b2-0a00-18f600000000
application/pdf
endstream
endobj
1 0 obj
<>
endobj
67 0 obj
<>/ProcSet[/PDF/Text/ImageC]/XObject<>>>/Type/Page>>
endobj
6 0 obj
<>/Font<>/ProcSet[/PDF/Text/ImageB/ImageC/ImageI]/XObject<>>>/Type/Page>>
endobj
12 0 obj
<>/ProcSet[/PDF/Text/ImageB]/XObject<>>>/Type/Page>>
endobj
18 0 obj
<>/ProcSet[/PDF/Text/ImageB]/XObject<>>>/Type/Page>>
endobj
24 0 obj
<>/ProcSet[/PDF/Text/ImageB]/XObject<>>>/Type/Page>>
endobj
30 0 obj
<>/ProcSet[/PDF/Text/ImageB]/XObject<>>>/Type/Page>>
endobj
36 0 obj
<>/ProcSet[/PDF/Text/ImageB]/XObject<>>>/Type/Page>>
endobj
42 0 obj
<>/ProcSet[/PDF/Text/ImageB]/XObject<>>>/Type/Page>>
endobj
48 0 obj
<>/ProcSet[/PDF/Text/ImageB]/XObject<>>>/Type/Page>>
endobj
88 0 obj
[90 0 R]
endobj
89 0 obj
<>stream
q
533.76 0 0 749.4 0 0 cm
/Im0 Do
Q
BT
/T1_0 1 Tf
8 0 0 8 319.39984 737 Tm
( on April 24, 2013)Tj
0 0 1 rg
-13.44897 0 Td
(rsta.royalsocietypublishing.org)Tj
0 g
-8.11399 0 Td
(Downloaded from )Tj
ET
endstream
endobj
46 0 obj
<>]/Filter[/CCITTFaxDecode]/Height 6245/Length 88473/Subtype/Image/Type/XObject/Width 4448>>stream
\!32sXz23$f|q*y:/telDT)ڼ3'`-͌Y:greܐV!!b+#Y"(dr`D0tCLD DNrUE;hhC!E8>Y֬Z (AhXr{ )!tDq|C({|'Ek!8a5sMN#@aѠ:A. 84rd(z!ß8PPP!M @}7*K!+jvv( d(0(0D-`EY+t.cMma[aow>
F=-ml =>D[a4C6]6p7Aw}^ V'[i5utiowt
'{o߰L5_ݵoVwa'wwMunaTVow۵w˲nj}wu{{KVäz][6.?xuoinWu~z.[ë:}CM7.]}zkL:[wW[߽=oҿUN_|E_U}
M5u_$;S.OBPy< _[ >-|/mwF9
l.VTtŲ"J͆>R}Ʒv$!o}U{w T~mV3kR7I4Ȝh{}WB
aUG3za~_KOb~G᷎ w<7OJh{_idh7_8o7&^êT~$3dZx'mh#wKZ1!{x'jߤ_}KVu=uV?Nj0^uVzTӫҰa֗AKMk~ᄑ U۷} jt_ҪҴdtGJ[mֻ{zU
HKmtKikVT}֖o/kݥ$Ai*Jꮛ }
* [A&Maa鄛=xv[VA0aPAM%ݥ0DEV0r4
ͨf"$s U4aK a5
Fsd\3D~? b`4
(D&Cԣ&0dl1
pt3b!&th2 ވ*x]vQa3^U>A2\2Ǻ[9OU|DDD097Jq"PH6g*L 8]`(gN3PBhCADDD\DDDDC".VDƌ{<˃DDDDZeDDDDDDDF"?ZY.ɺVZ!vd\K(bh&u@Mё;;X̫FV&dH2ĬgxHA LX ̖` ̔&IL>UA2+jL&vSN34OM}zʂAޮk"WTnL}0Bh.?TACA>>?~? |=vp:Ni߿wN?wi{M{J;^MWa0Tl,
"9
rTdj2
"{2 k*Io+vStjutK+tRޚ[uinםi^wu_J*Rh:_FD%$WoJR;%1ם8Tf] 9$RN5r"kn(*)j"c\*'&ʁ$"d* CPl@DFEç,勑28Α9/Rb#rI"LR3@P `Q0 4P"y `)T*&*N)̅FA@pj3@P.U -c\& "3AɆ`a22AALd$LCLȄGAAdq (I# hX& *iv.$n*K0!г3XD)E@sbd`a0L&gT*&N0AȀs lPN*@
L#aT203X!#,!0DH.&"bRJf(N!"va4-4OCAL"8jI(a4ӵD-EB0]2PEI i;BNAh ""(3BA
{`
4A r@!(A
A% kn4waU}r6A8, A"(M40}4 o
L(T@5@
o"6A8-'L J5dxL";hhboayl /aaMa[Tj
A''`{ "A7YOX D6A8-AvTtOiYN$vxO
ǢcI5l^æm& h";miC &6fGm&ؒMAMeBf*M.>}"@al̎N
Bf*Ai:AIMVt!ю
Da
zpa]pO]S% a6@Jpb2pMlzwELA&kpSi Aa)k 0xA PDBA~1[60ޚU}6/vM+5
];=# ݅A§+aV M>ij!݄a~[?]m~MSN֓{tޓMSモixK}^{UpuAM>-^^n:XtiKvtOW[^V~=?NL'{{Tծ/AݭVSZk־-SQwWvu]կmUi/kӰUAC8'P?!zo?KI*֒w~;p2V;V#EWˆUG
C-Ym#'-U6ZJIn?A/ǽhm5/IZOkȁ]//\w~%JJ
P~ȕ$~]r(zA_n4#N^39CGURt,B'%B"7]OY ]J?O
KдW@l]pDoB*wD>Tzh"7-l]]DopOIWz]8IU]UuwAi?_U'IIתA4æ ]w_o^4[ OM[ aIKvRon/zڷj^@k[~{.7_ozu}atI}at$a[{US_zwJv?I~ӷzk]{[ݯK{n.֗못I=pȃ/{mNR wZ[o ,D;맮J~}.}]k
ݮK^յuKW_iu[_߷KN_{=kQVF_y]Y!?kVZ!E{[_]]$[MkW U[~*KSͧ-SUkw~{O/q~ݑ VĕnzVD$ wzuUWA_am__I7~*U=9jڮSI~WZ[fg;i%Kno_n/
$m-k-{;U7s{WKfsٜ.l$iZ
-7 ^}i-{[[?~ku-M6[#@sRkk}[U4
1*+^I[JaT-I=;_m%n>l4N.N0jV",i < ]M
0E.Sa
v0[Va]
0Jh4DS0KB[H>:MH;aM)Vua}Uy,vPl0Et`҆ ! aQ`X8
.!
I!VvLB4P!R:jI2荌/;7L"
2btA) RN* ;J?;B(hI"0N k `ؒ"HSAHoe@0D90r_"J61"A"'BRPD@&VYAB$L
"C2aFFUh2t 28L7yXE80Uxfb$0L p< AI"3pLS8$r a0@D<8`.d\t0_&Ad-
4\SCRLH8pLXxA0L34T"fA0AʡmC>*M jFAij0{OD^j#BdP)rtEudpk
$84&a4`0O!@uA,AVp"
QA0H!æi&"TEAOI'\XM40DH
"-4; xT2 HzZi&SxAe'nk R$
vIA7դTD8IGFt
6ADPTH Di6iAOi i46II; aDt^"xPn A@<"X] 6nphtc848DXM1%~I: P a'
a0
D[TlÐAJC}7%wKSzN">V
P?_"m8OiR.B*mb+u]7 CAbxanz'YKW(pw@"DxE#V#q]}b߸[-Pj<:W-_E]}گ]JzW]pޤr(rn-H]W.'Nz-_koK!v_M>y R![h0jaa D =?
/ߑWnatA-_AHP i}?B^BRB/'_=c1aS/߂T0M[]E7݄Fw"uuU-.д*l6;>moP"o@~tp~?Dom5]|^R^W=]?m/ ^XH?I'A{K[߿z n/~
l70z[[
.ۮޗj%Ajڷ_w}_
ރ|=.A~_tEAoޮu]uנZ~oMҧak./^]uKav67aw_iww8A~[Z?&դui}UK_W%%?z]:]9*FkK4hn^-]/mO_wx_i5PKKۮU_ߪ_U]R}[_Vw0뮝]u}RIa_jN}:kmjkdmuZ\wY.ȅ_%u"$2W!6n_/h:jrm*k
za ߛ@ݮ][V}+-߶o\7}O]owKפo5^VkiZտW~ڽ{{ki~
oUm>U[g_j^Ӻh'I]l+>kn/iZn?V-WjmgvWWj۫[ -U}>M҆}P+a5amU
+M;Mwjl_Od~WMn}Z
M[JI$OOM0+0:'`VCŸKTNl0
[!+aUn
i,6 d}{
0E-#qA8`[a;M8a v:AB0C6Am/z`UHfzLS±\0bжtCalfqL<t$:h
Dn)#PӤA`PImB"7x#CB N&;b!
a(}M l0"PVxa(zl4`D(h.);a(5Xh$a!C
ᅈP pzPlI cӍ&AN´b&Tж<Պn0XL2:F8a &%ҦB $VǧS0ӄ
6F9uBa0?l&<L&1 L c0
UVj2:Ka@`8 kO%(`S
6wxkiV >ba
&U"4@&A.`fxaU5i;װI1Xi<1
1'"gsgAr "2Cٞl$gO L&5T>&IKy!G&A7LbAc]B*&a T`E74b",ha\ ,DI
4,0L&2ЙlT&k8GLa3hL(r(p/
ІH{3q2HtULA]B0i0ABB"B""",a$SB-DDDDD* 0!h2BC8SH"A,iH̜,10@\0Tr(r2B"""Bӈ&8&ss!Ą6uëb@!4"""vB""-CkIkUީ~U
H/-[AֵETPh¥XC0J#YSDTd$k-LiBDd5dn2A ٝ E!!dG2+2UVTȸ%'L)DG-#T-UTЌȂ7(2 [{~Cd4MaCd+8>dx%"ppl#v
ᅦ/t
n-a$l$u|#+Gv
u_d6À%j~w]YZ[_nu]w뾩[KǦz'H!WNm.%BSܛjL:⠌⬂6@`h9e1P\UKc.v ]I@3#*ae@a;Ȁ!@R4Meic#l psS%FnN"r*qF%)
QANdj!kPF9*rD/Yڨ.S@\0L ϼ5!aq0&G a0,0aCa A8`0&Jx9
Jq00L#'DGGLʶP*
0D(J"8ADx tX@@HR:& `!ŒA``R:I:Adq氅|&Y4- &@FSPL
"0B<@d`u l!%Y@!J+``@E DI3P`# A!^e8pLftH eCۤfd#!b4χL&`@pD[h^!"4Dz4D"3
<(A0W2L ig}xN#
aR(#A|&AƄh^L(D(QGa!B-$H7Z.6"x@A-"9A-tBpzay!
KAy"ѨA!\04DK[i9/:nX~xM;H.D^hX 6
#'7D4E tP
+d"=h&C 8j!
N ehn6澩6Fbb
MD8K6( " .+I6%A ~a|=DAȰ H7ZlACv.]?4( !;h4:,Ppa"B7[pl6nJ!҆OH7Acv$ê
T6oNl =:MCa$ rApI$NKPz
ӆA~M6um҆D℟ puMmuN4n;XAvit1WMVoP۲z6Mi6
ik&ڐCl'A$+uzp(%o ŧIWv&8UkCiviاza%[aa#J}wNWSJ0o+^
WWtNz]=_UTAtmZ|$i{ma%ai;n=MWW
ƭ}&ZUq6uM.ƶzl*j]&KPÅFMJۄӤii0UZT[n{knoWj+}խ%VJ _V [_ O@k7ZLgˆtڵ-*ڵ^09
0dp{ )G.ƞXX4+a|4%!iB{uXJ[HGv#C_Z q֫0bUi/Ӆ}{Oȕ,U"Uroۥ[UD3D8lu" Շպr(}%h9c=WCJOX_g,r uuJ \DK^7_}Һ4DF[uUo'_m%1kwui>Ku^@#uaK<ARv/I2n_}% NKuw]$]W}/p_ՇX_&3
_IɎ$Kl>ւ%Ңfl"7K_nҥ}X[zvK_._U .n넽+]O@kj __J-[mhwz%UoI:K{Uz" _TxnH>K}[U߭V@WA+/RKUA}^;iXZn]zn@]_'Bm+kuz+Aح{pzZWR_A_Wo_JK_Km%VK[k}_]/mJzۮ K}:_uA'N\%Km_Ur]zҫ]{^]NbOZ}}lwx%W~u}^U_U]&ׯD8wkiu}%^F]R\-m]"'t?Z=_Z^!}P/^MקKKo_֗/+cnlolRSh_^!kA6U.)[uV}OOKvOkڤzMN^:w_powkӻ]O'r2uխ[jҭ{J9]뮨5A6yT?7ki[a~_aUXkL'h7ۭ]]WUm4Zikul0i5an]u_L*no* o
v+ͫ5ӵg
0vպWo?7?UTһU\kj`t3@똘`5xm+u[[I vV~U!L0 :pa(au
u@
>҆aiյl'iXU! Ul*# 9ګպV/ lh
uUmoM
aTO* :^']7@b6P10 l `*a(y6)6f8vMP 0
lI0@m "ma S/7h`PE"!2!LD0#
EƁ^nDmXjAILpº
0< tئ GlTD&"][&b :aQx2'h&ӠhI`b6 ;Dn)I)и:
hT00H4t#1·aB6A+\Bba[< ;x`05M4,<0fa6lV b~; BNTJV ا _imMvB]DUlEvVi(2kiqç}0au.<@S^&DXUXw
;?# &;X0L/mc22}5J
8a ǃD2.i^Lq3U*e`I5$:L0hFh0&DDHL i"0- PL=IrAHE2(sP3eL&SD&<0Af{n03P
0`Ⱥ!\a8&pT""Be id-2c`!DCD0 AZ10ALe0`PDDDYPpB8 h0Dma`Mg e q XTЈ!
m~տaa*w8Ws Yؚuu`='R$DT!)X - BMNN $ICtBA PD@i\àDRB3q6
hJ
'i_
D,.wti81AԐv7IyPAMkKuNM!lBD!
&7 0[m(n^NqIL.{JWa pwqꇮ7%v o(ҶuBGMiZگU_~K]u߮K.h-ah T]
JK*]}]K.TJK{zw iݥC,0!&խ~E?ׄZK{okk}Iq׆d_um[l"o[m+_[[P>l*ڑ<`UVuC2&p( hC(N4%l"10a0X6)X0Κ˅t6`4-0Q a K``ICPA&!STh,uA
Ph\1Ai~XP0aATўT"jl5-3a0[VDZeDE!0A#D;B
l+8 aG궓jI]_]}+Ip%wA$ am>$B$nd @Cq,b?Zpe"3h8B8ae4AT͔(D&atӺ^[ZOzrPzq`¾0v`
V?|5oi1ك.f/gc";kL#'"b&25=W$F\TGd6(9@cM.xUlnAr<re \A՛A Ldv`˙GLd&E314Ps)dC. 41+=8G Du慄,NHX@ivr$s[lB"'\pᚢNF pa4-8D"D29k8݄Bds`q
oA5iվCLw!]"8AbB; Ap`҃B0d<"NadH%aB%!&Aa[ԉ,4tAB,& 6STp@FaB0'ǻA#ACI
!РA6
,8L#A!Aa&cIdžDNm߶A;p
npaA_t6&aV}'ۺ_?Ҵo ?[?w~^I?}l=Iꕷ^֝kNUP^۾VW^&nmM?ӧ}uvM,wu_];kW='kW{~\^s;ӺW&ߖ?a|(">7Z,}B3ձ_sG0^/}*U]9?7KˎMw>{{a֞/>7.uktl?Ҹn֒MwIj*vJۥ_~Io^C֫I /
a6_7-zˈzj(?wKJٰ_봫KV]⿇֯q%c_[R#o_c7K|I
7Ȱ{+GC
ԉ}?Cz;>zȰ{+ `Nȱ[5
5?
oz]>l;k
V^_[jpn{j4I7[:v?V*viګW$aa h40AUL.B@KU
mۤۿ 7NaD
apA<ZZAf0 *E8L A
h
Ԑ@¶I)
cPB̆pB v$Aal/ԡ V$pj
0aR
pFM
Ag80g
JC9&tm 0$A0:UYtxB)0]ޝ{8!Bb
6A(UMC@MkPgBCy,rC6!x;Zkܜ
#SNz"td>'vD0]C8^!g4
Љqc67p I974Z^A渎渎x0o&T7\çd'ЉQ
Lg6U" Ͱ9B""""0hD
Ef$DIjXbC2"" `8Wa
d"#;!ABU p'3IN0J5
Gb;M8rˢҘK >;#!8exgbI5N@X2)vdM3"kU=8r!Bhiއh4iW׆w;a^~O
``M/ׯvYIW2Q6epe'炼oPE$HU"7>tcuIU!ItꖯQzMKuI-^&_kmz1R{zmmx^@/Ho(]WI|DjrTEᢢDRM';fiIATڭ_b'XiUVWKî^꒭|PuIQ
QUGTmWS_VלLi+ZKQ=D1<4 j ^aekNDV)GL nN=_zXM$4
}WK6m&n[ 1&>ILtXNU-)P/GbMi34BUڠ8DeK}0 i:ezGR"LB&0W`A&b #JoChVT+##%*( &%@WCDC*C B"]RΝ=n!G͊2{UD3l0C2rbhD^#_]z 'pAu-Af ,&>&:،DGq >RA(9foY5Yh&30AZUBwz%w6
!i V+@XaoǭhI:;ao]}~vu*]tWޗGdؤY&&QV[5:'2!w&D6H
N̑\@#'
C @A
-#LUbh֎,KjMș;F0@62LT&c:fls a$
tP=M*#2L&VB*;$!&`BE`= e2.A0(L0! "ph^,~"-IH @<0DNdZ>3RPƳȀ^fΩ cC !8g&a45AO>*̠$A'4av!+a'"n 4BDEv! aa0 5@ϋa@JNꃃ@@Xh4]174KD_mL a"xul;xxA4Ë́"pktmj">=0Pv@EilM|"~m[arA;b߾timULSNIwiձ>u&k#aXIAul1Xj+nt^~u}KV0ҴtڮҼ>>6WOkw߫]6m~zmkMki7ޕ;KoN^U_/釦Ն;BW:;_+_cj/n_Ȏv_W__]Bֿov3/G___V.R~E%KkOkwvj5/UWHu7Z^oU__qMuo{oK^NЮ۪D/
]]_wi?]-'
_^B5u_U7I/~Cz]Dp~+ZE:}#둉^kWTW-Sk~]AxWOVNzwKvoN[ilN{U5[>I5MPeխtKlGmKl=_ڦiY}իjvJph%4lH&ViZMwt[]7p;V`ҵNڢ5a#8rC@a NSz^wUzwU Bnl `h+PAC+
NIpV0ܷ@ I[I =ckiH J7Th4
q. ڃ C&Ճ#b6"0D1i2p) #8atBLWA`5Xa$¶5I0AAzAV)iwLBdSB`b,5n
6#^إa$ĸ
aVvUV t8j0AA;ɧL([0ADp@ a4CpLB[fAB
5W]͆@'qjn"u $
-0"9ڬ `0`B xdT5
0MC,D03D03n